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Gene: Aqp5 MGI:106215

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Not currently registered for phenotyping at IMPC

Phenotyping is currently not planned for a knockout strain of this gene.

Gene Summary

Name:
aquaporin 5
Synonyms:
N/A

Order Alleles

IMPC Data Collections

  • No Body Weight Data
  • No Embryo Imaging Data
  • No Viability Data

IMPC Phenotype Summary

Significant
Not Significant
Not tested

View all our phenotype data below

Phenotypes

The IMPC applies a panel of phenotyping screens to characterise single-gene knockout mice by comparison to wild types. Click on the different tabs to visualise significant phenotypes identified by the IMPC, as well as all data that was measured.

lacZ Expression

Expression data not available

Associated Images

Images submitted by IMPC centres for a selection of procedures. Each set of images is available to view in our image comparator.

Phenotype associated images not available

Human diseases caused by Aqp5 mutations

The analysis uses data from IMPC, along with published data on other mouse mutants, in comparison to human disease reports in OMIM, Orphanet, and DECIPHER.

Phenotype comparisons summarize the similarity of mouse phenotypes with human disease phenotypes.

The table below shows human diseases associated to Aqp5 by orthology or direct annotation.

Disease Similarity of
phenotypes		 Matching phenotypes Source
Palmoplantar Keratoderma, Bothnian Type
OMIM:600231
Non-Epidermolytic Palmoplantar Keratoderma
ORPHA:2337

The table below shows human diseases predicted to be associated to Aqp5 by phenotypic similarity.

Disease Similarity of
phenotypes		 Matching phenotypes Source
Keratoendotheliitis Fugax Hereditaria
Keratitis, Epiphora, Conjunctival hyperemia, Opacification of the corneal stroma OMIM:148200
Corneal Dystrophy, Meesmann, 1
Punctate opacification of the cornea, Corneal dystrophy, Epiphora OMIM:122100
Alacrima, Congenital, Autosomal Dominant
Lacrimal gland hypoplasia, Punctate corneal epithelial erosions, Alacrima, Decreased lacrimation OMIM:103420
Corneal Dystrophy, Meesmann, 2
Epiphora, Recurrent corneal erosions OMIM:618767
Epithelial Recurrent Erosion Dystrophy
Corneal erosion, Epiphora OMIM:122400
Limbal Stem Cell Deficiency
Lacrimation abnormality, Opacification of the corneal epithelium, Generalized opacification of th... ORPHA:171673
Alacrima, Congenital, Autosomal Recessive
Punctate corneal epithelial erosions, Alacrima OMIM:601549
Vernal Keratoconjunctivitis
Lacrimation abnormality, Abnormal cornea morphology, Abnormal conjunctiva morphology, Punctate ke... ORPHA:70476
Herpes Simplex Virus Stromal Keratitis
Keratitis, Deep anterior chamber, Herpetiform corneal ulceration, Corneal stromal edema, Corneal ... ORPHA:137599
Epithelial Recurrent Erosion Dystrophy
Subepithelial corneal opacities, Corneal dystrophy, Irregular astigmatism, Corneal scarring, Kera... ORPHA:293381
Corneal Dystrophy, Posterior Polymorphous, 1
Polymorphous posterior corneal dystrophy, Iris atrophy, Corneal opacity, Band keratopathy, Uveal ... OMIM:122000
Ring Dermoid Of Cornea
Corneal astigmatism, Abnormal corneal limbus morphology, Abnormal cornea morphology, Abnormal con... OMIM:180550
Intellectual Developmental Disorder, X-Linked, Syndromic 17
Anisocoria, Alacrima OMIM:300858
Gelatinous Drop-Like Corneal Dystrophy
Subepithelial corneal opacities, Central opacification of the cornea, Conjunctival amyloidosis, C... ORPHA:98957
Isolated Congenital Alacrima
Keratitis, Corneal erosion, Lacrimal gland hypoplasia, Conjunctivitis, Alacrima ORPHA:91416
Neuropathy, Hereditary Sensory And Autonomic, Type Viii
Corneal ulceration, Corneal scarring, Decreased lacrimation OMIM:616488
Hyperuricemia, Infantile, With Abnormal Behavior And Normal Hypoxanthine Guanine Phosphoribosyltransferase
Alacrima OMIM:240000
Neurotrophic Keratopathy
Lacrimation abnormality, Corneal scarring, Corneal stromal edema, Astigmatism, Recurrent corneal ... ORPHA:137596
Posterior Polymorphous Corneal Dystrophy
Increased corneal curvature, Corneal opacity, Lacrimation abnormality, Uveal ectropion, Abnormal ... ORPHA:98973
Dacryocystitis-Osteopoikilosis Syndrome
Lacrimation abnormality ORPHA:1562
Keratitis-Ichthyosis-Deafness Syndrome, Autosomal Recessive
Keratoconus, Conjunctivitis, Decreased lacrimation OMIM:242150
Lacrimal Duct Defect
Dacryocystocele, Conjunctivitis, Epiphora OMIM:149700
Keratosis Pilaris Atrophicans
Epiphora OMIM:604093
Paroxysmal Extreme Pain Disorder
Lacrimation abnormality OMIM:167400
Contractures-Ectodermal Dysplasia-Cleft Lip/Palate Syndrome
Lacrimation abnormality ORPHA:1484
Hydroa Vacciniforme
Keratitis, Epiphora ORPHA:330058
Classic Progressive Supranuclear Palsy Syndrome
Conjunctival hyperemia, Decreased lacrimation ORPHA:240071
Neuropathy, Hereditary Sensory And Autonomic, Type I, With Cough And Gastroesophageal Reflux
Alacrima OMIM:608088
Laryngotracheoesophageal Cleft
Neonatal respiratory distress, Dyspnea, Stridor, Cough, Impaired oropharyngeal swallow response, ... ORPHA:2004
Growth Hormone Deficiency, Isolated Partial
Postnatal growth retardation, Short stature, Decreased response to growth hormone stimulation test OMIM:615925
Intellectual Disability-Alacrima-Achalasia Syndrome
Anisocoria, Alacrima ORPHA:289483
Lacrimoauriculodentodigital Syndrome 2
Conjunctivitis, Alacrima OMIM:620192
Corneal Dystrophy, Fuchs Endothelial, 1
Corneal dystrophy, Corneal stromal edema, Corneal guttata, Corneal degeneration, Descemet Membran... OMIM:136800
Otofaciocervical Syndrome 2, With T-Cell Deficiency
Conjunctivitis, Alacrima, Lacrimal duct stenosis OMIM:615560
Cockayne Syndrome Type 1
Cryptorchidism, Cataract, Conjunctivitis, Decreased lacrimation ORPHA:90321
Arrhinia-Choanal Atresia-Microphthalmia Syndrome
Lacrimation abnormality ORPHA:1135
Intestinal Botulism
Respiratory insufficiency due to muscle weakness, Dyspnea, Diarrhea, Xerostomia, Dysphagia ORPHA:178481
Familial Dysautonomia
Corneal opacity, Abnormal pupil morphology, Corneal erosion, Heterochromia iridis, Alacrima ORPHA:1764
Hypohidrotic Ectodermal Dysplasia-Hypothyroidism-Ciliary Dyskinesia Syndrome
Lacrimation abnormality ORPHA:1882
Chronic Pneumonitis Of Infancy
Respiratory distress, Intercostal retractions, Reduced forced vital capacity, Tachypnea, Hypoxemi... ORPHA:91359
Frontonasal Dysplasia 2
Bilateral cryptorchidism, Decreased lacrimation OMIM:613451
Central Hypoventilation Syndrome, Congenital, 3
Apnea, Central hypoventilation, Chronic constipation, Respiratory failure, Episodic vomiting OMIM:619483
Peripheral Demyelinating Neuropathy, Central Dysmyelination, Waardenburg Syndrome, And Hirschsprung Disease
Cryptorchidism, Heterochromia iridis, Alacrima, Decreased lacrimation OMIM:609136
Neuropathy, Hereditary Sensory And Autonomic, Type Iii
Alacrima, Corneal ulceration, Recurrent corneal erosions OMIM:223900
Nasolacrimal Duct Cyst
Chronic irritative conjunctivitis, Corneal astigmatism, Nasolacrimal duct obstruction, Dacryocyst... ORPHA:141083
Graft Versus Host Disease
Decreased lacrimation ORPHA:39812
Blepharonasofacial Malformation Syndrome
Cryptorchidism, Lacrimation abnormality ORPHA:1252
Stuve-Wiedemann Syndrome 1
Opacification of the corneal stroma, Decreased lacrimation OMIM:601559
Sunct Syndrome
Conjunctival hyperemia, Increased tear production, Epiphora ORPHA:57145
Achalasia-Addisonianism-Alacrima Syndrome
Anisocoria, Alacrima OMIM:231550
Trisomy 8P
Cryptorchidism, Astigmatism, Aplasia/Hypoplasia of the gallbladder, Annular pancreas, Heterochrom... ORPHA:264450
Haddad Syndrome
Small for gestational age, Central hypoventilation, Polyhydramnios, Breathing dysregulation, Gast... ORPHA:99803
Cockayne Syndrome
Cataract, Band keratopathy, Cryptorchidism, Developmental cataract, Keratoconjunctivitis sicca, A... ORPHA:191
Neurodevelopmental Disorder With Seizures, Hypotonia, And Brain Imaging Abnormalities
Neonatal respiratory distress, Decreased response to growth hormone stimulation test, Polyhydramn... OMIM:618922
Cleft Larynx, Posterior
Aspiration OMIM:215800
Paroxysmal Hemicrania
Conjunctival hyperemia, Epiphora ORPHA:157835
Cockayne Syndrome B
Cryptorchidism, Developmental cataract, Microcornea, Hypoplasia of the iris, Opacification of the... OMIM:133540
Obesity-Hypoventilation Syndrome
Hypoventilation, Obesity OMIM:257500
Focal Facial Dermal Dysplasia Type Iii
Lacrimation abnormality ORPHA:1807
Alacrimia-Choreoathetosis-Liver Dysfunction Syndrome
Corneal neovascularization, Corneal scarring, Decreased lacrimation ORPHA:404454
Blepharophimosis-Ptosis-Epicanthus Inversus Syndrome Plus
Streak ovary, Elevated circulating luteinizing hormone level, Polycystic ovaries, Lacrimal gland ... ORPHA:572333
Nasopalpebral Lipoma-Coloboma Syndrome
Conjunctival hyperemia, Epiphora OMIM:167730
Lacrimoauriculodentodigital Syndrome 3
Alacrima OMIM:620193
Corneal Endothelial Dystrophy
Abnormal Descemet membrane morphology, Increased corneal thickness, Corneal dystrophy, Opacificat... OMIM:217700
Yao Syndrome
Asthma, Xerostomia, Diarrhea, Weight loss OMIM:617321
Helix Syndrome
Hyperparathyroidism, Alacrima OMIM:617671
Persistent Hyperplastic Primary Vitreous
Cataract, Corneal opacity, Leukocoria, Developmental cataract, Microcornea, Buphthalmos, Shallow ... ORPHA:91495
Cockayne Syndrome A
Cryptorchidism, Cataract, Opacification of the corneal stroma, Decreased lacrimation OMIM:216400
Iatrogenic Botulism
Dyspnea, Constipation, Xerostomia, Dysphagia ORPHA:254509
Alacrima, Achalasia, And Impaired Intellectual Development Syndrome
Anisocoria, Alacrima OMIM:615510
Chondroectodermal Dysplasia With Night Blindness
Epiphora ORPHA:319195
Kilquist Syndrome
Alacrima OMIM:619080
Peripheral Demyelinating Neuropathy-Central Dysmyelinating Leukodystrophy-Waardenburg Syndrome-Hirschsprung Disease
Heterochromia iridis, Decreased lacrimation ORPHA:163746
Neuropathy, Hereditary Sensory And Autonomic, Type Vi
Alacrima, Corneal scarring OMIM:614653
Multiple Congenital Anomalies-Hypotonia-Seizures Syndrome 4
Astigmatism, Alacrima OMIM:618548
Branchiootorenal Syndrome 1
Gustatory lacrimation, Lacrimation abnormality, Euthyroid goiter, Lacrimal duct stenosis OMIM:113650
Waardenburg Syndrome Type 1
Lacrimation abnormality, Heterochromia iridis ORPHA:894
Inhalational Botulism
Dyspnea, Diarrhea, Xerostomia, Constipation ORPHA:254504
Nasopalpebral Lipoma-Coloboma Syndrome
Conjunctival hyperemia, Cataract, Corneal opacity, Epiphora ORPHA:2399
Foodborne Botulism
Respiratory insufficiency due to muscle weakness, Diarrhea, Xerostomia, Cardiorespiratory arrest,... ORPHA:228371
Muscular Dystrophy, Limb-Girdle, Autosomal Recessive 18
Cataract, Alacrima OMIM:615356
Botulism
Diarrhea, Xerostomia, Respiratory insufficiency, Constipation, Dysphagia ORPHA:1267
Achalasia, Familial Esophageal
Achalasia, Xerostomia OMIM:200400
Amyotrophic Lateral Sclerosis
Dyspnea, Respiratory failure, Xerostomia, Abnormal respiratory system physiology ORPHA:803
Congenital Disorder Of Glycosylation, Type Im
Aspiration, Diarrhea, Failure to thrive, Vomiting OMIM:610768
Congenital Disorder Of Deglycosylation 1
Corneal ulceration, Corneal opacity, Alacrima OMIM:615273
Developmental And Epileptic Encephalopathy 38
Aspiration, Gastroesophageal reflux OMIM:617020
Neurodevelopmental Disorder With Dysmorphic Facies, Impaired Speech, And Hypotonia
Cryptorchidism, Alacrima OMIM:619005
Mucoepithelial Dysplasia, Hereditary
Cataract, Keratoconjunctivitis, Opacification of the corneal stroma, Corneal neovascularization, ... OMIM:158310
Antisynthetase Syndrome
Edema, Cough, Xerostomia, Respiratory insufficiency, Dysphagia, Pulmonary arterial hypertension ORPHA:81
Erythermalgia, Primary
Diarrhea, Constipation, Xerostomia, Hyperhidrosis OMIM:133020
Deeah Syndrome
Decreased response to growth hormone stimulation test, Anterior pituitary hypoplasia, Cryptorchid... OMIM:619004
Hyperostosis Cranialis Interna
Epiphora OMIM:144755
Hyperekplexia 1
Aspiration, Apnea OMIM:149400
Rapid-Onset Childhood Obesity-Hypothalamic Dysfunction-Hypoventilation-Autonomic Dysregulation Syndrome
Decreased response to growth hormone stimulation test, Adrenocorticotropic hormone deficiency, Go... ORPHA:293987
Acrocraniofacial Dysostosis
Lacrimation abnormality ORPHA:949
Cone-Rod Dystrophy 10
Epiphora OMIM:610283
Dyskeratosis Congenita, Autosomal Recessive 1
Pterygium, Nasolacrimal duct obstruction, Epiphora OMIM:224230
Infant Botulism
Bowel incontinence, Respiratory insufficiency due to muscle weakness, Dyspnea, Xerostomia, Consti... ORPHA:178478
Laminin Subunit Alpha 2-Related Congenital Muscular Dystrophy
Hypoventilation, Respiratory insufficiency, Dysphagia, Pulmonary arterial hypertension, Respirato... ORPHA:258
Pontine Tegmental Cap Dysplasia
Aspiration, Failure to thrive, Dysphagia OMIM:614688
Lacrimoauriculodentodigital Syndrome 1
Absence of Stensen duct, Nasolacrimal duct obstruction, Lacrimal gland hypoplasia, Conjunctivitis... OMIM:149730
Waardenburg Syndrome
Lacrimation abnormality, Heterochromia iridis ORPHA:3440
Amyotrophic Lateral Sclerosis 21
Respiratory insufficiency due to muscle weakness, Aspiration, Dysphagia OMIM:606070
Reynolds Syndrome
Xerostomia, Respiratory insufficiency, Gastroesophageal reflux, Dysphagia, Ascites ORPHA:779
Alternating Hemiplegia Of Childhood
Respiratory distress, Apnea, Oral-pharyngeal dysphagia, Diarrhea, Dehydration, Hyperhidrosis, Vom... ORPHA:2131
Vocal Cord And Pharyngeal Distal Myopathy
Respiratory insufficiency due to muscle weakness, Aspiration, Dysphagia ORPHA:600
Postaxial Polydactyly-Dental And Vertebral Anomalies Syndrome
Lacrimation abnormality ORPHA:2916
Cone-Rod Dystrophy 8
Epiphora OMIM:605549
Esophageal Atresia
Respiratory distress, Small for gestational age, Failure to thrive in infancy, Polyhydramnios, Ep... ORPHA:1199
Johanson-Blizzard Syndrome
Lacrimation abnormality ORPHA:2315
Tay-Sachs Disease
Aspiration OMIM:272800
Sjogren Syndrome
Xerostomia OMIM:270150
Oculopharyngodistal Myopathy 1
Respiratory distress, Hypercapnia, Reduced forced vital capacity, Respiratory insufficiency due t... OMIM:164310
Central Hypoventilation Syndrome, Congenital, 1
Hypoventilation, Apnea, Central hypoventilation, Hypercapnia, Hyperhidrosis, Hypoxemia, Chronic c... OMIM:209880
Carcinoid Syndrome
Epiphora ORPHA:100093
Postorgasmic Illness Syndrome
Xerostomia, Hyperhidrosis ORPHA:279947
Silver-Russell Syndrome Due To Maternal Uniparental Disomy Of Chromosome 7
Small for gestational age, Decreased response to growth hormone stimulation test, Hyperhidrosis, ... ORPHA:96182
Dyskeratosis Congenita, Autosomal Recessive 6
Epiphora OMIM:616353
Eec Syndrome
Lacrimation abnormality, Decreased response to growth hormone stimulation test, Keratitis, Cornea... ORPHA:1896
Halperin-Birk Syndrome
Aspiration, Pseudobulbar paralysis, Failure to thrive, Gastroesophageal reflux OMIM:618651
Schinzel-Giedion Syndrome
Annular pancreas, Streak ovary, Alacrima ORPHA:798
Spinocerebellar Ataxia Type 8
Aspiration, Dysphagia ORPHA:98760
Diffuse Cutaneous Systemic Sclerosis
Dyspnea, Xerostomia, Gastroesophageal reflux, Dysphagia, Pulmonary arterial hypertension ORPHA:220393
Adiposis Dolorosa
Diarrhea, Xerostomia, Obesity, Constipation, Hypothyroidism ORPHA:36397
Hypohidrotic Ectodermal Dysplasia
Sinusitis, Xerostomia, Hypohidrosis, Cough, Failure to thrive ORPHA:238468
Lissencephaly Type 1 Due To Doublecortin Gene Mutation
Aspiration, Dysphagia ORPHA:2148
Al Amyloidosis
Gastrointestinal hemorrhage, Gastroparesis, Dyspnea, Nonproductive cough, Xerostomia, Weight loss... ORPHA:85443
Oculocerebrorenal Syndrome Of Lowe
Hyperparathyroidism, Cataract, Corneal opacity, Lacrimation abnormality, Cryptorchidism, Abnormal... ORPHA:534
Gm2-Gangliosidosis, Ab Variant
Aspiration OMIM:272750
Prune Belly Syndrome
Xerostomia, Oligohydramnios OMIM:100100
Mixed Connective Tissue Disease
Gastrointestinal hemorrhage, Gastritis, Dyspnea, Xerostomia, Joint swelling, Gastroesophageal ref... ORPHA:809
Rett Syndrome, Congenital Variant
Aspiration, Constipation, Gastroesophageal reflux OMIM:613454
Lacrimoauriculodentodigital Syndrome
Increased corneal thickness, Cryptorchidism, Keratoconjunctivitis, Keratoconjunctivitis sicca, Re... ORPHA:2363
Polyposis, Skin Pigmentation, Alopecia, And Fingernail Changes
Cachexia, Diarrhea, Xerostomia, Hematochezia, Vomiting OMIM:175500
Dyskeratosis Congenita, Autosomal Dominant 3
Cryptorchidism, Epiphora OMIM:613990
Dyskeratosis Congenita, Digenic
Decreased testicular size, Epiphora OMIM:620040
Ramos-Arroyo Syndrome
Respiratory distress, Xerostomia, Chronic constipation, Decreased body weight, Severe failure to ... ORPHA:1051
Neuromuscular Oculoauditory Syndrome
Respiratory distress, Aspiration OMIM:618733
Opitz Gbbb Syndrome
Aspiration, Gastroesophageal reflux, Dysphagia OMIM:300000
Burning Mouth Syndrome
Xerostomia ORPHA:353253
Adnp Syndrome
Respiratory distress, Oral-pharyngeal dysphagia, Chronic constipation, Truncal obesity, Gastroeso... ORPHA:404448
Williams Syndrome
Flat cornea, Corneal opacity, Cataract, Lacrimation abnormality, Cryptorchidism, Blue irides, Pol... ORPHA:904
Ectodermal Dysplasia 11B, Hypohidrotic/Hair/Tooth Type, Autosomal Recessive
Anhidrosis, Rhinitis, Xerostomia OMIM:614941
Charge Syndrome
Cryptorchidism, Anterior hypopituitarism, Lacrimation abnormality, Iris coloboma ORPHA:138
Aplasia Of Lacrimal And Salivary Glands
Xerostomia OMIM:180920
Developmental And Epileptic Encephalopathy 100
Aspiration, Gastroesophageal reflux, Dysphagia, Chronic constipation OMIM:619777
Amoebiasis Due To Free-Living Amoebae
Corneal perforation, Conjunctival hyperemia, Corneal ulceration, Increased tear production ORPHA:68
Dyskeratosis Congenita, X-Linked
Cataract, Cryptorchidism, Conjunctivitis, Pterygium, Decreased testicular size, Epiphora OMIM:305000
Bilateral Perisylvian Polymicrogyria
Apnea, Pseudobulbar paralysis, Gastroesophageal reflux, Dysphagia, Aspiration ORPHA:98889
Chronic Graft Versus Host Disease
Dyspnea, Wheezing, Pneumothorax, Xerostomia, Bronchiectasis, Dysphagia, Diarrhea, Weight loss, Ga... ORPHA:99921
Sarcoidosis, Susceptibility To, 1
Abnormal salivary gland morphology, Enlarged lacrimal glands, Epiphora OMIM:181000
Stüve-Wiedemann Syndrome
Lacrimation abnormality, Ectopic thyroid ORPHA:3206
Ogden Syndrome
Pulmonary edema, Apnea, Lymphedema, Diarrhea, Pulmonary arterial hypertension, Restrictive ventil... OMIM:300855
Lambert-Eaton Myasthenic Syndrome
Constipation, Xerostomia, Hypohidrosis ORPHA:43393
Stevens-Johnson Syndrome/Toxic Epidermal Necrolysis Spectrum
Symblepharon, Keratitis, Corneal erosion, Keratoconjunctivitis sicca, Conjunctivitis, Increased t... ORPHA:95455
Neurodevelopmental Disorder With Hypotonia And Dysmorphic Facies
Neonatal respiratory distress, Apnea, Decreased response to growth hormone stimulation test, Feta... OMIM:619503
Rubinstein-Taybi Syndrome Due To Ep300 Haploinsufficiency
Pneumonia, Polyhydramnios, Asthma, Obesity, Gastroesophageal reflux, Constipation, Aspiration, Fa... ORPHA:353284
Rubinstein-Taybi Syndrome Due To Crebbp Mutations
Pneumonia, Polyhydramnios, Asthma, Obesity, Gastroesophageal reflux, Constipation, Aspiration, Fa... ORPHA:353277
Igg4-Related Dacryoadenitis And Sialadenitis
Palpebral edema, Facial edema, Xerostomia, Thyroiditis, Weight loss ORPHA:79078
Igg4-Related Submandibular Gland Disease
Facial edema, Periorbital edema, Prostatitis, Xerostomia ORPHA:449432
Blau Syndrome
Dyspnea, Joint swelling, Xerostomia, Pulmonary arterial hypertension ORPHA:90340
Non-Epidermolytic Palmoplantar Keratoderma
ORPHA:2337
Palmoplantar Keratoderma, Bothnian Type
OMIM:600231

Histopathology

Summary table of phenotypes displayed during the Histopathology procedure which are considered significant. Full histopathology data table, including submitted images, can be accessed by clicking any row in this table.

There is no histopathology data for Aqp5

IMPC related publications

The table below lists publications which used either products generated by the IMPC or data produced by the phenotyping efforts of the IMPC. These publications have also been associated to Aqp5.

No publications found that use IMPC mice or data for Aqp5.

Order Mouse and ES Cells

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MGI Allele Allele Type Produced
Aqp5tm1aNarl KO first allele (reporter-tagged insertion with conditional potential) Mice, ES Cells
Aqp5em3(IMPC)H Intra-exon deletion Mice
Aqp5em2(IMPC)H Intra-exon deletion Mice

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